{"id":3953,"date":"2012-09-18T22:19:38","date_gmt":"2012-09-19T04:19:38","guid":{"rendered":"http:\/\/earbirding.com\/blog\/?p=3953"},"modified":"2012-09-18T22:21:05","modified_gmt":"2012-09-19T04:21:05","slug":"the-two-part-calls-of-empidonax","status":"publish","type":"post","link":"http:\/\/earbirding.com\/blog\/archives\/3953","title":{"rendered":"The &#8220;Two-part Calls&#8221; of Empidonax"},"content":{"rendered":"<figure id=\"attachment_3960\" aria-describedby=\"caption-attachment-3960\" style=\"width: 300px\" class=\"wp-caption alignright\"><a href=\"http:\/\/www.flickr.com\/photos\/bryanto\/4722753004\/\"><img loading=\"lazy\" decoding=\"async\" class=\"size-medium wp-image-3960\" title=\"HAFL_photo\" src=\"http:\/\/earbirding.com\/blog\/wp-content\/uploads\/2012\/09\/HAFL_photo-300x225.jpg\" alt=\"\" width=\"300\" height=\"225\" srcset=\"http:\/\/earbirding.com\/blog\/wp-content\/uploads\/2012\/09\/HAFL_photo-300x225.jpg 300w, http:\/\/earbirding.com\/blog\/wp-content\/uploads\/2012\/09\/HAFL_photo-150x112.jpg 150w, http:\/\/earbirding.com\/blog\/wp-content\/uploads\/2012\/09\/HAFL_photo-400x300.jpg 400w, http:\/\/earbirding.com\/blog\/wp-content\/uploads\/2012\/09\/HAFL_photo.jpg 1024w\" sizes=\"auto, (max-width: 300px) 100vw, 300px\" \/><\/a><figcaption id=\"caption-attachment-3960\" class=\"wp-caption-text\">If only it would sing! Probable Hammond&#39;s Flycatcher, Wyoming, 6\/17\/2010. Photo by Bryant Olsen (CC-by-nc-2.0)<\/figcaption><\/figure>\n<p><em>Empidonax<\/em> flycatchers are tough to identify by sight.\u00a0 Every birder knows it.\u00a0 They&#8217;re the classic bugaboo of North American bird identification.\u00a0 That&#8217;s why every field guide mentions the importance of listening to their voices.<\/p>\n<p>But Empids make a lot of sounds.\u00a0 Forget about learning &#8220;<em>the<\/em> song&#8221; and &#8220;<em>the<\/em> call.&#8221;\u00a0 Most Empids have repertoires of 6-8 different songs and calls. Some species, such as Pacific-slope and Cordilleran, have a <a title=\"http:\/\/earbirding.com\/blog\/archives\/2996\" href=\"http:\/\/earbirding.com\/blog\/archives\/2996\" target=\"_blank\">dawn song<\/a> that&#8217;s different from anything they say during the day.\u00a0 Several, including Least, Yellow-bellied, Hammond&#8217;s, and Dusky, have complex, rarely-heard flight songs. The species with the largest vocabulary appears to be Acadian Flycatcher, which has all of the above types of song plus another type, sometimes called the &#8220;<a title=\"http:\/\/www.xeno-canto.org\/33587\" href=\"http:\/\/www.xeno-canto.org\/33587\" target=\"_blank\">evening song<\/a>,&#8221; which is the most complex of all.\u00a0 (It may or may not be fully separate from the flight song.)<\/p>\n<p>Today I&#8217;m going to be talking about a class of Empid vocalizations that don&#8217;t get much press. I&#8217;ll call them &#8220;Two-part Calls&#8221; since they don&#8217;t have an official name.\u00a0 Based on their similarity, the &#8220;two-part calls&#8221; appear to be homologous &#8212; that is, evolutionarily equivalent, all descended from the same calling behavior of a common ancestor.<\/p>\n<p>As far as I know, three species of Empids give these calls. In one species, the two-part call is familiar enough to be mentioned in field guides, at least. The two-part call of the second species is described only in the scientific literature.\u00a0 And that of the third is, as far as I know, being described in this blog post for the first time.<\/p>\n<h4>Dusky Flycatcher: &#8220;Du-hic&#8221;<\/h4>\n<p>This call is mentioned in the <em>Sibley Guide to Birds<\/em>, Kenn Kaufmann&#8217;s <em>Advanced Birding<\/em>, and other well-researched field guides.\u00a0 The &#8220;du&#8221; part of the call is nearly monotone, and the &#8220;hic&#8221; is shorter and slightly higher.\u00a0 As you can hear in the following examples, the sequence is often more like &#8220;du, du, du, du, du-hic&#8221;.\u00a0 Sometimes the &#8220;hic&#8221; notes will be given without a &#8220;du,&#8221; or after other &#8220;hics.&#8221;<\/p>\n<table>\n<tbody>\n<tr>\n<td><div class=\"iframe-wrapper\">\n  <iframe src=\"http:\/\/www.xeno-canto.org\/embed.php?XC=12206&#038;simple=0\" frameborder=\"0\" style=\"height:220px;width:400px;\">Please upgrade your browser<\/iframe>\n<\/div><\/td>\n<td><div class=\"iframe-wrapper\">\n  <iframe src=\"http:\/\/www.xeno-canto.org\/embed.php?XC=65848&#038;simple=0\" frameborder=\"0\" style=\"height:220px;width:400px;\">Please upgrade your browser<\/iframe>\n<\/div><\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<p>This call may be given primarily by males, but I&#8217;m not certain of that. It&#8217;s given throughout the breeding season, but especially in long bouts at dawn and dusk during the early summer, prior to egg laying.<\/p>\n<h4>Hammond&#8217;s Flycatcher: &#8220;Peer-pewit&#8221;<\/h4>\n<p>Quite similar to Dusky&#8217;s &#8220;du-hic&#8221; but not mentioned in any field guides that I know, this call was first described in the scientific literature by James Sedgwick in 1975. Sedgwick called it the &#8220;k-lear whee-zee&#8221; call, but I think &#8220;peer-pewit&#8221; is a better transliteration. The &#8220;peer&#8221; note is slightly more downslurred than Dusky&#8217;s &#8220;du,&#8221; and the &#8220;pewit&#8221; is higher, longer, and much more distinctly two-syllabled than Dusky&#8217;s &#8220;hic.&#8221;\u00a0 It&#8217;s given in similar situations to the &#8220;du-hic,&#8221; though it&#8217;s apparently\u00a0 more likely to be heard later in the breeding season, after egg laying.<\/p>\n<div class=\"iframe-wrapper\">\n  <iframe src=\"http:\/\/www.xeno-canto.org\/embed.php?XC=14431&#038;simple=0\" frameborder=\"0\" style=\"height:220px;width:400px;\">Please upgrade your browser<\/iframe>\n<\/div>\n<h4>Yellow-bellied Flycatcher: &#8220;Whit-beert&#8221;<\/h4>\n<p>I stumbled across this vocalization in a good recording by Randy Little from Herkimer County, New York, which appears to be the only such recording in existence. The call is clearly related to the &#8220;du-hic&#8221; and &#8220;peer-pewit&#8221; of the western species, but instead of a drawn-out whistle, the first note is an emphatic &#8220;whit!&#8221; that resembles a more explosive version of the &#8220;whit&#8221; calls of other Empids. The second note is again an up-down-up trace on the spectrogram, rather like the Hammond&#8217;s &#8220;pewit&#8221; note.<\/p>\n<figure id=\"attachment_3967\" aria-describedby=\"caption-attachment-3967\" style=\"width: 614px\" class=\"wp-caption alignnone\"><a href=\"http:\/\/earbirding.com\/blog\/wp-content\/uploads\/2012\/09\/YBFL_whit-beert.jpg\"><img loading=\"lazy\" decoding=\"async\" class=\"size-full wp-image-3967\" title=\"YBFL_whit-beert\" src=\"http:\/\/earbirding.com\/blog\/wp-content\/uploads\/2012\/09\/YBFL_whit-beert.jpg\" alt=\"\" width=\"614\" height=\"181\" srcset=\"http:\/\/earbirding.com\/blog\/wp-content\/uploads\/2012\/09\/YBFL_whit-beert.jpg 614w, http:\/\/earbirding.com\/blog\/wp-content\/uploads\/2012\/09\/YBFL_whit-beert-300x88.jpg 300w, http:\/\/earbirding.com\/blog\/wp-content\/uploads\/2012\/09\/YBFL_whit-beert-150x44.jpg 150w, http:\/\/earbirding.com\/blog\/wp-content\/uploads\/2012\/09\/YBFL_whit-beert-400x117.jpg 400w\" sizes=\"auto, (max-width: 614px) 100vw, 614px\" \/><\/a><figcaption id=\"caption-attachment-3967\" class=\"wp-caption-text\">Yellow-bellied Flycatcher &quot;whit-beert&quot; call, Woodhull Lake, New York, 5\/30\/1998. ML catalog #106901<\/figcaption><\/figure>\n<p>(Click <a title=\"http:\/\/macaulaylibrary.org\/audio\/106901\" href=\"http:\/\/macaulaylibrary.org\/audio\/106901\" target=\"_blank\">here<\/a> to listen to the original recording on the Macaulay Library website.)<\/p>\n<p>I cannot find any definite mention of the &#8220;whit-beert&#8221; call of Yellow-bellied Flycatcher in any published source.\u00a0 The only possible reference I&#8217;ve found is this brief statement in the Birds of North America account:<\/p>\n<blockquote><p>One variation of <em>Tu-Wee<\/em> Call is of longer duration, described as <em>thoo weep eh, thoo weep eh<\/em>, or <em>she weeps sir<\/em> (<a href=\"https:\/\/cuvpn.colorado.edu\/bna\/species\/566\/articles\/biblio\/,DanaInfo=bna.birds.cornell.edu+bib035\">Hausman 1946<\/a>), or <em>pea-wayk-pea-wa<\/em><em>yk<\/em> (Dr. Hoy in <a href=\"https:\/\/cuvpn.colorado.edu\/bna\/species\/566\/articles\/biblio\/,DanaInfo=bna.birds.cornell.edu+bib022\">Forbush 1927<\/a>) and may actually represent call of different function.<\/p><\/blockquote>\n<p>That&#8217;s it.\u00a0 Not much to go on.<\/p>\n<p>So we&#8217;re left with more questions than answers.\u00a0 Do other Empids give &#8220;two-part calls&#8221;?\u00a0 What functions do they serve in each species?\u00a0 Are they ever given by migrants?\u00a0 Are they very rare in the Yellow-bellied Flycatcher, or merely under-reported?<\/p>\n<p>I&#8217;d be interested to hear anybody else&#8217;s experience with this type of call in Empids.<\/p>\n","protected":false},"excerpt":{"rendered":"<p>As far as I know, three species of Empids give these calls. In one species, the two-part call is familiar enough to be mentioned in field guides, at least. The two-part call of the second species is described only in the scientific literature.  And that of the third is, as far as I know, being described in this blog post for the first time.<\/p>\n","protected":false},"author":1,"featured_media":0,"comment_status":"open","ping_status":"open","sticky":false,"template":"","format":"standard","meta":{"footnotes":""},"categories":[103,4],"tags":[302,305,301,303,300,304],"class_list":["post-3953","post","type-post","status-publish","format-standard","hentry","category-behavior","category-id","tag-dusky-flycatcher","tag-empidonax-flaviventris","tag-empidonax-hammondii","tag-empidonax-oberholseri","tag-hammonds-flycatcher","tag-yellow-bellied-flycatcher"],"_links":{"self":[{"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/posts\/3953","targetHints":{"allow":["GET"]}}],"collection":[{"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/posts"}],"about":[{"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/types\/post"}],"author":[{"embeddable":true,"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/users\/1"}],"replies":[{"embeddable":true,"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/comments?post=3953"}],"version-history":[{"count":14,"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/posts\/3953\/revisions"}],"predecessor-version":[{"id":3971,"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/posts\/3953\/revisions\/3971"}],"wp:attachment":[{"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/media?parent=3953"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/categories?post=3953"},{"taxonomy":"post_tag","embeddable":true,"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/tags?post=3953"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}