{"id":4180,"date":"2013-01-19T13:33:57","date_gmt":"2013-01-19T19:33:57","guid":{"rendered":"http:\/\/earbirding.com\/blog\/?p=4180"},"modified":"2013-01-19T13:33:57","modified_gmt":"2013-01-19T19:33:57","slug":"splitting-the-golden-fronted-woodpecker","status":"publish","type":"post","link":"http:\/\/earbirding.com\/blog\/archives\/4180","title":{"rendered":"Splitting the Golden-fronted Woodpecker"},"content":{"rendered":"<p><a href=\"http:\/\/schmoker.org\/BirdPics\/Woodpeckers.html\"><img loading=\"lazy\" decoding=\"async\" class=\"alignright size-medium wp-image-4181\" title=\"GFWO1\" src=\"http:\/\/earbirding.com\/blog\/wp-content\/uploads\/2013\/01\/GFWO1-300x223.jpg\" alt=\"Golden-fronted Woodpecker, McAllen, Texas, April 2004. Photo courtesy of Bill Schmoker.\" width=\"300\" height=\"223\" srcset=\"http:\/\/earbirding.com\/blog\/wp-content\/uploads\/2013\/01\/GFWO1-300x223.jpg 300w, http:\/\/earbirding.com\/blog\/wp-content\/uploads\/2013\/01\/GFWO1-150x111.jpg 150w, http:\/\/earbirding.com\/blog\/wp-content\/uploads\/2013\/01\/GFWO1-400x298.jpg 400w, http:\/\/earbirding.com\/blog\/wp-content\/uploads\/2013\/01\/GFWO1.jpg 576w\" sizes=\"auto, (max-width: 300px) 100vw, 300px\" \/><\/a>The taxonomy of the Golden-fronted Woodpecker and its relatives has been causing headaches for over a century.\u00a0 Not only does the species display complex geographic variation in plumage, but where ranges meet, it hybridizes with the closely related Gila Woodpecker to the northwest, the Red-bellied Woodpecker to the northeast, and the Hoffman&#8217;s Woodpecker to the south, which in turn sometimes hybridizes with the Red-crowned Woodpecker in Costa Rica.\u00a0 The result is a mind-bending mosaic of similar-looking woodpeckers stretching from Ontario to Venezuela.<\/p>\n<p>Current taxonomy draws species boundaries despite the hybridization, in part due to differences in vocalizations.\u00a0 For example, <a title=\"http:\/\/xeno-canto.org\/102885\" href=\"http:\/\/xeno-canto.org\/102885\" target=\"_blank\">Red-bellied<\/a> and <a title=\"http:\/\/xeno-canto.org\/7198\" href=\"http:\/\/xeno-canto.org\/7198\" target=\"_blank\">Hoffmann&#8217;s<\/a> Woodpeckers both give a long, churring rattle call that Golden-fronted lacks.\u00a0 Vocal differences probably help the woodpeckers keep each other straight in areas where ranges meet, making them more likely than not to choose a mate of their own taxon, keeping the level of gene flow low and maintaining the boundary between the species in the long term.<\/p>\n<p>As I was going through the Golden-fronted Woodpecker recordings in my collection, some of which are from Texas and some of which are from Mexico, I realized that at least one call in the Golden-fronted repertoire varies geographically.\u00a0 Birds in southern Texas and northern Mexico give a rough, single-syllabled &#8220;gaf&#8221; call rather like that of the Red-bellied Woodpecker:<\/p>\n<table>\n<tbody>\n<tr>\n<td><div class=\"iframe-wrapper\">\n  <iframe src=\"http:\/\/www.xeno-canto.org\/embed.php?XC=5773&#038;simple=0\" frameborder=\"0\" style=\"height:220px;width:400px;\">Please upgrade your browser<\/iframe>\n<\/div><\/td>\n<td><div class=\"iframe-wrapper\">\n  <iframe src=\"http:\/\/www.xeno-canto.org\/embed.php?XC=109193&#038;simple=0\" frameborder=\"0\" style=\"height:220px;width:400px;\">Please upgrade your browser<\/iframe>\n<\/div><\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<p>But birds from Veracruz south give a distinctive, two-syllabled &#8220;CHUCK-a&#8221;:<\/p>\n<table>\n<tbody>\n<tr>\n<td><div class=\"iframe-wrapper\">\n  <iframe src=\"http:\/\/www.xeno-canto.org\/embed.php?XC=118429&#038;simple=0\" frameborder=\"0\" style=\"height:220px;width:400px;\">Please upgrade your browser<\/iframe>\n<\/div><\/td>\n<td><div class=\"iframe-wrapper\">\n  <iframe src=\"http:\/\/www.xeno-canto.org\/embed.php?XC=118430&#038;simple=0\" frameborder=\"0\" style=\"height:220px;width:400px;\">Please upgrade your browser<\/iframe>\n<\/div><\/td>\n<\/tr>\n<tr>\n<td><div class=\"iframe-wrapper\">\n  <iframe src=\"http:\/\/www.xeno-canto.org\/embed.php?XC=33955&#038;simple=0\" frameborder=\"0\" style=\"height:220px;width:400px;\">Please upgrade your browser<\/iframe>\n<\/div><\/td>\n<td><\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<p>These differences hold up in every recording I&#8217;ve managed to find of this call type.\u00a0 The single-note &#8220;gaf&#8221; can be heard at least as far north as the <a title=\"http:\/\/blb.biosci.ohio-state.edu\/LongData.asp?RecordingID=11890\" href=\"http:\/\/blb.biosci.ohio-state.edu\/LongData.asp?RecordingID=11890\" target=\"_blank\">Texas Panhandle<\/a> and as far south as <a title=\"http:\/\/macaulaylibrary.org\/audio\/23117\" href=\"http:\/\/macaulaylibrary.org\/audio\/23117\" target=\"_blank\">San Luis Potosi<\/a>.\u00a0 The double-noted &#8220;CHUCK-a&#8221; has been recorded not only in <a title=\"http:\/\/macaulaylibrary.org\/audio\/23102\" href=\"http:\/\/macaulaylibrary.org\/audio\/23102\" target=\"_blank\">Veracruz<\/a> but also in <a title=\"http:\/\/macaulaylibrary.org\/audio\/137684\" href=\"http:\/\/macaulaylibrary.org\/audio\/137684\" target=\"_blank\">Guatemala<\/a> and <a title=\"http:\/\/macaulaylibrary.org\/audio\/20912\" href=\"http:\/\/macaulaylibrary.org\/audio\/20912\" target=\"_blank\">El Salvador<\/a>.<\/p>\n<p>It turns out that these differences almost exactly mirror the geographic boundaries between two major subspecies groups.\u00a0 The &#8220;<em>aurifrons<\/em> group&#8221; is the &#8220;classic&#8221; Golden-fronted Woodpecker of Texas and northern Mexico.\u00a0 The double-noted birds appear to correspond to the &#8220;<em>santacruzi<\/em>&#8221; group, which is found from Veracruz all the way south to Honduras (including the &#8220;<em>polygrammus<\/em>&#8221; group of southern Pacific-slope Mexico).\u00a0 It&#8217;s worth noting that the subspecies <em>dubius<\/em> from the Yucatan Peninsula apparently gives a slightly different version of the two-syllabled call, more of a \u201cchuck-trrr\u201d [<a title=\"http:\/\/macaulaylibrary.org\/audio\/23110\" href=\"http:\/\/macaulaylibrary.org\/audio\/23110\" target=\"_blank\">1<\/a> <a title=\"http:\/\/macaulaylibrary.org\/audio\/23111\" href=\"http:\/\/macaulaylibrary.org\/audio\/23111\" target=\"_blank\">2<\/a>], sometimes shortened to a single-noted \u201cchuck\u201d rather like the call of <em>aurifrons<\/em> [<a title=\"http:\/\/macaulaylibrary.org\/audio\/103346\" href=\"http:\/\/macaulaylibrary.org\/audio\/103346\" target=\"_blank\">3<\/a>].<\/p>\n<p>A <a title=\"http:\/\/www.researchgate.net\/publication\/231217705_Molecular_Systematics_of_the_Red_-Bellied_and_Golden-Fronted_Wood_peckers\/file\/79e4150660fc8484f9.pdf\" href=\"http:\/\/www.researchgate.net\/publication\/231217705_Molecular_Systematics_of_the_Red_-Bellied_and_Golden-Fronted_Wood_peckers\/file\/79e4150660fc8484f9.pdf\" target=\"_blank\">2009 genetic analysis<\/a> by Erick Garc\u00eda-Trejo and colleagues found that the &#8220;<em>aurifrons<\/em>&#8221; group and the &#8220;<em>santacruzi<\/em>&#8221; group were genetically distinct, with northern <em>aurifrons<\/em> more closely related to the Red-bellied Woodpecker, <em>Melanerpes carolinus<\/em>, than to southern birds.\u00a0 As announced in <a title=\"http:\/\/www.aba.org\/birding\/v42n4p28.pdf\" href=\"http:\/\/www.aba.org\/birding\/v42n4p28.pdf\" target=\"_blank\"><em>Birding<\/em> magazine<\/a> and elsewhere, this set the stage for a formal split of the species by the AOU into Golden-fronted Woodpecker (<em>M. aurifrons<\/em>) and Velasquez&#8217;s Woodpecker (<em>M. santacruzi<\/em>).\u00a0 However, nobody has apparently submitted a formal proposal to the North American Checklist Committee yet &#8212; so I took it upon myself to do that.\u00a0 Sometime later this year, we&#8217;ll find out what they say!<\/p>\n","protected":false},"excerpt":{"rendered":"<p> At least one call in the Golden-fronted Woodpecker&#8217;s repertoire varies geographically, corresponding to the geographic boundaries between two major subspecies groups that probably deserve species rank.<\/p>\n","protected":false},"author":1,"featured_media":0,"comment_status":"open","ping_status":"open","sticky":false,"template":"","format":"standard","meta":{"footnotes":""},"categories":[16],"tags":[320,321,323,322],"class_list":["post-4180","post","type-post","status-publish","format-standard","hentry","category-taxonomy","tag-golden-crowned-woodpecker","tag-melanerpes-aurifrons","tag-melanerpes-santacruzi","tag-velasquezs-woodpecker"],"_links":{"self":[{"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/posts\/4180","targetHints":{"allow":["GET"]}}],"collection":[{"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/posts"}],"about":[{"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/types\/post"}],"author":[{"embeddable":true,"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/users\/1"}],"replies":[{"embeddable":true,"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/comments?post=4180"}],"version-history":[{"count":10,"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/posts\/4180\/revisions"}],"predecessor-version":[{"id":4192,"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/posts\/4180\/revisions\/4192"}],"wp:attachment":[{"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/media?parent=4180"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/categories?post=4180"},{"taxonomy":"post_tag","embeddable":true,"href":"http:\/\/earbirding.com\/blog\/wp-json\/wp\/v2\/tags?post=4180"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}